Ecologist Daniel Simberloff has investigated the role of habitat fragmentation in species extinction. He documented that when large continuous forests are divided into smaller isolated patches, certain bird species disappear even when the total area of remaining forest equals or exceeds their minimum territorial requirements. Studies tracking individual birds revealed that these species rarely cross open areas between fragments, effectively rendering the isolated patches inaccessible. Moreover, populations confined to single fragments exhibited reduced genetic diversity over successive generations compared to populations in continuous habitat. Mathematical models incorporating these dispersal limitations and genetic factors predict higher extinction probabilities for fragmented populations than for equally sized continuous populations, even when controlling for total habitat area. This research indicates that ____